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If you interact with him without it, you won't understand what he's saying – only gibberish – and the quest won't progress.

Progression: For the quest to start, you will need to equip the Spore Shroom Charm before talking to him.

Location: You will find Mister Mushroom standing right where Cornifer used to sell his map in the fungal wastes after defeating the 3 Dreamers.

Mister Mushroom is a mysterious entity first located in the fungal wastes and part of the mushroom "Hivemind" This can be seen as he seems to be talking to other entities including the Hungry Knight a character for a Team Cherry game of the same title. Mister Mushroom is an NPC in Hollow Knight. He will start appearing in various locations of the map after defeating the 3 Dreamers.